The Epaulette Shark, Hemiscyllium ocellatum, subadult. Image: Scott W. Michael.

Genus Hemiscyllium

By Scott W. Michael

Excerpt from CORAL Magazine, September/October 2021



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Heinrich, D.D., Rummer, J.L., Morash, A.J., Watson, S.A., Simpfendorfer, C.A., Heupel, M.R. and Munday, P.L., 2014. A product of its environment: the epaulette shark (Hemiscyllium ocellatum) exhibits physiological tolerance to elevated environmental CO2. Conservation Physiology, 2(1), p.cou047.

Heupel, M.R. and Bennett, M.B., 1998. Observations on the diet and feeding habits of the epaulette shark, Hemiscyllium ocellatum (Bonnaterre), on Heron Island Reef, Great Barrier Reef, Australia. Marine and Freshwater Research49(7), pp.753-756.

Heupel, M.R. and Bennett, M.B., 1999. The occurrence, distribution and pathology associated with gnathiid isopod larvae infecting the epaulette shark, Hemiscyllium ocellatumInternational Journal for Parasitology29(2), pp.321-330.

Heupel, M. R. and M. B. Bennett. 2007. Estimating Abundance of Reef-Dwelling Sharks: A Case Study of the Epaulette Shark, Hemiscyllium ocellatum (Elasmobranchii: Hemiscyllidae) 1. Pacific Science, 61(3), pp.383-394.

Jutan, Y., Retraubun, A.S.W., Khouw, A.S., Nikijuluw, V.P.H. and Latumaerissa, H.S., 2019, October. The food composition of Halmahera walking shark (Hemiscyllium halmahera). In IOP Conference Series: Earth and Environmental Science (Vol. 339, No. 1, p. 012007). IOP Publishing.

Jutan, Y., Retraubun, A.S.W., Khouw, A.S., Nikiju-luw, V.P.H. and Pattikawa, J.A., 2018. Study on the population of Halmahera walking shark (Hemiscyllium halmahera) in Kao Bay, North Maluku, Indonesia. Interna-tional Journal of Fisheries and Aquatic Studies, 6(4), pp.36-41.

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Peach, M.B., 2002. Rheotaxis by epaulette sharks, Hemiscyllium ocellatum (Chondrichthyes: Hemiscylliidae), on a coral reef flat. Australian Journal of Zoology, 50(4), pp.407-414.

Soderstrom, V., Renshaw, G.M. and Nilsson, G.E., 1999. Brain blood flow and blood pressure during hypoxia in the epaulette shark Hemiscyllium ocellatum, a hypoxia-tolerant elasmobranch. Journal of experimental biology, 202(7), pp.829-835.

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Special thanks to Vanessa Sommers of Oceans Reefs and Aquariums ( for providing information about epaulette shark breeding and husbandry. 

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